RT Journal Article T1 Biogeografía histórica de las clavatoráceas (carófitas fósiles). Estado actual del conocimiento T1 Historical biogeography of Clavatoraceae (fossil charophytes). State of the art A1 Martín Closas, Carles A1 Serra Kiel, Josep AB The historical biogeography of Clavatoraceae (Mesozoic charophytes) is analysed on the basis of phylogeneticanalysis and the data available about its distribution. The most plesiomorphic and oldest representativesof this family occurred during the Late Jurassic and were scattered in a few localities of the central par t ofthe peri-Tethyan domain (Western Europe and Northern Africa) and the margins of the North Americanseaway (Morrison Fm). In the Neocomian, clavatoraceans were abundant in the peri-Tethyan region andexpanded eastwards to reach the Chinese basins in the Valanginian or Hauterivian. During Barremian andAptian the clavatoraceans reached their maximum diversity in the peri-Tethyan region and their maximumpaleogeographic extension worldwide. Thus, in the Early Barremian the family is recorded from southernAmerica, where it migrated probably through Northwest Africa. Also in the Aptian, it was also present inNorth America. Only a few species were able to reach a cosmopolitan distribution during Barremian andAptian. This is the case of the Atopochara trivolvis lineage. Some characters, such as the conjoint dispositionof gametangia are extremely significant to understand the worldwide migration of this species. In theAlbian the decline of the clavatoraceans began. After a significant gap in its fossil record, the latemostCretaceous clavatoraceans are relict forms (Atopochara trivolvis var. ulanensis) or belong to newly evolvedspecies of poorly known phylogenetic affinity (traditional Septorella). The history of Clavatoraceae largelyagrees with the asymmetric biogeographic model of López-Martínez (2003) YR 2006 FD 2006 LK http://hdl.handle.net/10272/8449 UL http://hdl.handle.net/10272/8449 LA spa DS Repositorio Institucional de la Universidad de Huelva RD 30 may 2026